Dragonfly wings are the complex result of multiple patterning systems interweaving in response to force flows and material properties. They consist of both honeycomb patterns which are flexible and exhibit membrane behavior and ladder-type patterns which are stiff and exhibit beam-like behavior. These patterns are characterized by their rule-based interaction in terms of cell density, cell shape, and cell depth, as well as other parameters affecting overall wing performance, such as out-of-plane pleating behavior and material distribution. This complex mineral skeleton is skinned with translucent cuticle which eliminates shear failure in-plane. The composite morphology of skeleton and skin is what ultimately generates wing performance.